Nature lovers held a vigil for a famous female grizzly bear that delighted visitors for decades in Wyoming’s Grand Teton National Park after she was struck and killed by a vehicle.

At 28 years old, Grizzly No. 399 was the oldest known reproducing female grizzly in the Yellowstone ecosystem.

During her nearly three decades roaming the snowy wilderness of the Teton Range, Grizzly No. 399 became a beloved mama bear with millions of parkgoers following her yearly adventures and her ever-growing family tree.

During her nearly three decades roaming the snowy wilderness of the Teton Range, Grizzly No. 399 became a beloved mama bear with millions of parkgoers following her yearly adventures and her ever-growing family tree.

Named for the tag affixed by researchers to her ear, No. 399 was the oldest-known reproducing female grizzly in the greater Yellowstone ecosystem